The latest imaging technologies mean that we can look underneath Titian's paintings.
The images reveal the faint outline of a redrawn arm, the ghost of a figure once on the right, now to the left.
Restorer Jill Dunkerton looks at how Titian adjusted some compositions as he worked to capture the moment of maximum impact.
Modern discoveries about Titian's 'poesie'
A look 'beneath' Titian's canvases reveals the tweaks and changes he made as he worked over four hundred years ago. Find out more with Restorer Jill Dunkerton.