The latest imaging technologies mean that we can look underneath Titian's paintings.
The images reveal the faint outline of a redrawn arm, the ghost of a figure once on the right, now to the left.
Restorer Jill Dunkerton looks at how Titian adjusted some compositions as he worked to capture the moment of maximum impact.
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A look 'beneath' Titian's canvases reveals the tweaks and changes he made as he worked over four hundred years ago. Find out more with Restorer Jill Dunkerton.